1. Georgia Cooperative Fish and Wildlife Research Unit, Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia, USA
2. Department of Forestry, Wildlife, and Fisheries, University of Tennessee, Knoxville, Tennessee, USA
3. U.S. Geological Survey, GA Cooperative Fish and Wildlife Research Unit, Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia, USA