1. Department of Fisheries, Wildlife and Conservation Biology, University of Minnesota, Minneapolis, Minnesota, USA
2. Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia, USA
3. Division of Fish and Wildlife, Minnesota Department of Natural Resources, Saint Paul, Minnesota,USA
4. U.S. Geological Survey, Minnesota Cooperative Fish and Wildlife Research Unit, Department of Fisheries, Wildlife and Conservation Biology, University of Minnesota, Minneapolis, Minnesota, USA
5. Department of Rangeland, Wildlife, and Fisheries Management, Texas A&M University, College Station, Texas, USA