1. a Laboratory of Protistology and Aquatic Ecology , Ghent University , Krijgslaan 281 S8, 9000 , Gent , Belgium
2. b Royal Botanic Garden , Edinburgh EH3 5LR, UK
3. c J. Craig Venter Institute , San Diego , California , 92121 , USA
4. d Aquatic Ecosystems , Institute for Food and Agricultural Research and Technology (IRTA) Crta Poble Nou s/n , 43540 , St Carles de la Rapita , Catalonia , Spain