1. The Marine Biological Association of the United Kingdom, The Laboratory, Citadel Hill, Plymouth PL1 2PB, UK
2. School of Ocean and Earth Science, University of Southampton, Southampton SO14 3ZH, UK
3. BRIDGE, School of Geographical Sciences, University of Bristol, University Road, Bristol BS8 1SS, UK
4. Plymouth Electron Microscopy Centre, Brunel, Drake Circus, Plymouth PL4 8AA, UK