1. Geum River Environment Research Center, National Institute of Environmental Research, Jiyongstreet, Okcheon gun, Chungcheongbukdo 29027, Republic of Korea
2. Graduate School of Fisheries Science & Environmental Studies, Nagasaki University, 1-14 Bunkyo-Machi, Nagasaki 852-8521, Japan
3. Metagenomics Research Group, National Research Institute of Fisheries Science, Fisheries Research Agency, 2-12-4 Fukuura, Kanazawa-Ku, Yokohama 236-8648, Japan
4. Laboratory of Environmental Biology, Department of Applied Aquabiology, National Fisheries University, 2-7-1 Nagata-Honmachi, Shimonoseki, Yamasguchi 759-6595, Japan
5. Jeju Center, Korea Basic Science Institute (KBSI), 213-4 Cheomdan-ro, Jeju City, Jeju Special Self-Governing Province 63309, Korea