1. Institute for Chemistry and Biology of the Marine Environment, Carl von Ossietzky University Oldenburg, Schleusenstraße 1, 26382 Wilhelmshaven, Germany
2. Department of Marine Science and Fisheries, Sultan Qaboos University, Al Khoud 123, P.O. Box 34, Muscat, Oman
3. Centre of Excellence in Marine Biotechnology, Sultan Qaboos University, Al Khoud 123, P.O. Box 50, Muscat, Oman