1. NIHR Nottingham Biomedical Research Centre, Nottingham, UK;
2. Department of Hearing Sciences, Division of Clinical Neuroscience, School of Medicine, University of Nottingham, Nottingham, UK;
3. Nottingham University Hospitals NHS Trust, Nottingham, UK;
4. Division of Cancer and Stem Cells, Academic Unit of Oncology, School of Medicine, University of Nottingham, Nottingham, UK