1. A.N. Severtsov Institute of Ecology and Evolution, Moscow, Russia
2. Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the RAS, Vladivostok, Russia
3. Pacific Salmons Department, Russian Federal Research Institute of Fisheries and Oceanography, Moscow, Russia
4. Departmant of Invertebrate Zoology, Faculty of Biology, Lomonosov Moscow State University, Moscow, Russia