1. Animal Health and Comparative Medicine, Institute of Biodiversity, University of Glasgow, Glasgow, Scotland, UK
2. Department of Biosciences, College of Life and Environmental Sciences, University of Exeter, Exeter, UK
3. School of Natural Sciences, University of Stirling, Stirling, UK
4. University Marine Biological Station, Millport, Isle of Cumbrae, Scotland, UK
5. School of Biological Sciences, University of Aberdeen, Aberdeen, UK