1. Ocean and Ecosystem Sciences Division, Maritimes Region, Department of Fisheries & Oceans, Bedford Institute of Oceanography, Dartmouth, Nova Scotia, Canada
2. Population Ecology Division, Maritimes Region, Department of Fisheries & Oceans, Bedford Institute of Oceanography, Dartmouth, Nova Scotia, Canada
3. Life Science Department, Los Angeles Pierce College, Woodland Hills, CA, USA
4. School of Aquatic and Fishery Sciences and Burke Museum of Natural History and Culture, University of Washington, Seattle, WA, USA