1. College of Agriculture and Food Sciences, King Faisal University, Hofuf, Saudi Arabia
2. Coastal and Offshore Resources Research Center, Fisheries Research Institute, Council of Agriculture, Kaohsiung, Taiwan
3. Centre of Advanced Study in Marine Biology, Annamalai University, Parangipettai, India
4. Center for Environment and Marine Studies, Research Institute, King Fahd University of Petroleum and Minerals, Dhahran, Saudi Arabia