1. Research Unit Ecosystems and Aquatics Resources (UR03AGRO1), Institut National Agronomique de Tunisie, University of Carthage, Tunis Mahrajéne, Tunisia
2. Marine and Coastal Protected Areas Department, Coastline Protection and Management Agency, Tunis Le Belvédère, Tunisia
3. Laboratorio de Biología Marina, Departamento de Zoología, Universidad de Sevilla, Sevilla, Spain