1. Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu, Sabah, Malaysia;
2. Faculty of Natural Science and Sustainability, University College Sabah Foundation, Sabah, Malaysia
3. Department of Zoology, Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, Sarawak, Malaysia
4. Department of Biodiversity and Conservation Biology, Biology Centre, Czech Academy of Sciences, Ceske Budejovice, Czech Republic