1. Laboratory of Apiculture and Social Insects, School of Life Sciences, University of Sussex, Brighton, UK
2. Department of Entomology and Acarology, Luiz de Queiroz College of Agriculture, University of São Paulo, Piracicaba, Brazil
3. Department of Zoology, Bioscience Institute, University of São Paulo, São Paulo, Brazil
4. California Department of Food & Agriculture, Plant Pest Diagnostics Center, Sacramento CA, USA
5. Honey Bee Research and Extension Laboratory, Department of Entomology and Nematology, University of Florida, Gainesville FL, USA