1. Department of Ecology, University of Innsbruck, Innsbruck, Austria
2. University of Natural Resources and Life Sciences, Department of Integrative Biology and Biodiversity Research, Institute of Integrative Conservation Research, Vienna, Austria
3. Institute of Bee Health, Vetsuisse Faculty, University of Bern, Bern, Switzerland
4. Agroscope, Swiss Bee Research Centre, Bern, Switzerland
5. CREA Research Centre for Agriculture and Environment, Bologna, Italy