1. a Department of Biological Sciences , National University of Singapore , 14 Science Drive 4, Singapore , 117543 , Republic of Singapore
2. b Center for Aquatic Conservation and Department of Biological Sciences , University of Notre Dame , Notre Dame , IN , 46556 , USA
3. c Marine Biodiversity and Biosecurity, National Institute of Water and Atmospheric Research , Private Bag 14901, Kilbirnie , Wellington , New Zealand
4. d Transdisciplinary Research Organization for Subtropical and Island Studies, University of the Ryukyus , Taketomi , Okinawa , 907-1541 , Japan
5. e Department of Biology , Universiti Brunei Darussalam, Jalan Tunku Link , Gadong , BE-1410 , Brunei Darussalam