1. a Aquatic Bioinvasions Research and Policy Institute, Environmental Sciences and Management, PO Box 751, Portland State University , Portland , Oregon , 97207-0751 , USA
2. b Smithsonian Environmental Research Center, Marine Invasions Research Laboratory, Romberg Tiburon Center for Environmental Studies , 3152 Paradise Drive, Tiburon , California , 94920 , USA
3. c Smithsonian Environmental Research Center, Marine Invasions Research Laboratory , PO Box 28, Edgewater , Maryland , 21037 , USA