1. Key Laboratory of Yak Breeding Engineering Gansu Province, Lanzhou Institute of Husbandry and Pharmaceutical Sciences, Chinese Academy of Agricultural Sciences, Lanzhou, China
2. Key Laboratory of Animal Genetics and Breeding on Tibetan Plateau, Ministry of Agriculture and Rural Affairs, Lanzhou, China
3. Life science and Engineering College, Northwest Minzu University, Lanzhou, China
4. Qilian County Veterinary Animal Husbandry Station, Qinghai, China
5. Huazhi Biotech Co. Ltd, Changsha, China