1. a Department of Animal Sciences , University of Illinois at Urbana-Champaign , Urbana , Illinois , USA
2. b CSIRO Livestock Industries, St Lucia , Queensland , Australia
3. c Department of Veterinary and Biomedical Sciences , University of Minnesota , St Paul , Minnesota , USA
4. d Institute for Genomic Biology, University of Illinois at Urbana-Champaign , Urbana , Illinois , USA