1. Department of Fisheries, Fisheries Research Institute (FRI), Glami Lemi, Titi, 71650 Jelebu, Negeri Sembilan, Malaysia
2. Department of Marine Science, Kulliyyah of Science, International Islamic University of Malaysia, Kuantan, Pahang, Malaysia
3. Department of Fisheries, Fisheries Research Institute (FRI), 11960 Batu Maung, Penang, Malaysia