1. Wildlife Conservation Research Unit, Recanati-Kaplan Centre, Department of Zoology, University of Oxford, Oxford, UK
2. CEFE, CNRS, University Montpellier, University Paul Valéry Montpellier, EPHE, IRD, Montpellier, France
3. Mammal Research Institute, Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa
4. Department of Computer Science, University of Oxford, Oxford, UK