1. a Department of Fisheries and Wildlife Sciences , Virginia Polytechnic Institute and State University , Blacksburg, VA 24061-0321, USA
2. d Horseshoe Crab Research Center , Virginia Polytechnic Institute and State University , Blacksburg, VA 24061-0321, USA
3. b National Marine Fisheries Service RTR Unit at Virginia Tech , 100 Cheatham Hall, Blacksburg, VA 24061-0321, USA
4. c Virginia-Maryland Regional College of Veterinary Medicine, Virginia Polytechnic Institute and State University , Blacksburg, VA 24061, USA