1. Department of Biodiversity Sciences, Graduate school of Social and Cultural Studies, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka 819–0395, Japan
2. National Museum of Nature and Science, 3-23-1, Hyakunin-cho, Shinjuku-ku, Tokyo 169–0073, Japan
3. Faculty of Bioresources, Mie University, 1577 Kurimamachiya-cyo, Tsu, Mie 514–8507, Japan
4. Marine World Umino-Nakamichi, 18–28, Saitozaki, Higashi-ku, Fukuoka, 811–0321, Japan