1. Faculty of Agriculture, Iwate University, Morioka, Iwate 020-8550, Japan
2. Tohoku Natural History Research Group, Morioka, Iwate 020-8550, Japan
3. Kyushu Environmental Evaluation Association, 1-10-1 Matsukadai, Higashi-ku, Fukuoka-shi, Fukuoka 813-0004, Japan
4. Iwate Prefectural Museum, 34 Aza-Matsuyashiki, Ueda, Morioka, Iwate 020-0102, Japan
5. Research and Education Centre for Natural Sciences, Keio University, Yokohama, Kanagawa 223-8521, Japan