1. Department of Biological Science, Faculty of Science, Nara Women's University, Nara 630-8506, Japan
2. Coastal Ecosystem Section, National Research Institute of Fisheries and Environment of Inland Sea, Maruishi 2-17-5, Hatsukaichi, Hiroshima 739-0452, Japan
3. Department of Civil Engineering, Tokushima University, Tokushima 770-8506, Japan