1. Department of Chemistry, Biology, and Marine Science, Faculty of Science, University of the Ryukyus, Okinawa 903-0213, Japan
2. Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790-8577, Japan
3. Joint Faculty of Veterinary Medicine, Yamaguchi University, Yamaguchi 753-8515, Japan
4. Miyagi Prefecture Fisheries Technology Institute, Ishinomaki, Miyagi 986-2135, Japan