1. Bioscience and Biotechnology Center, Nagoya University, Nagoya, Aichi 464-8601, Japan
2. Bioscience and Biotechnology Center, Nagoya University, Nagoya 464-8601, Japan
3. Yatomi Station, Aichi Fisheries Research Institute, Yatomi, Aichi 498-0017, Japan
4. Department of Marine Bioscience, Fukui Prefectural University, Obama, Fukui 917-0003, Japan