1. Department of Biology, Faculty of Sciences, Golestan University, Box 155, Gorgan, 39361-79142, Golestan, Iran
2. Department of Crop Productions Technology, Faculty of Agriculture, Higher Educational Complex of Saravan, Saravan 99516-34145, Iran
3. University of Gothenburg, Department of Biology and Environmental Sciences, Systematics and Biodiversity, Box 463, SE-405 30 Göteborg, Sweden