1. Sugashima Marine Biological Laboratory, Graduate School of Science, Nagoya University, Sugashima, Toba, Mie 517-0004, Japan
2. Misaki Marine Biological Station, School of Science, The University of Tokyo, Miura, Kanagawa 238-0225, Japan
3. Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo, Tokyo 113-0033, Japan
4. Faculty of Human Environmental Studies, Hiroshima Shudo University, Ozuka-Higashi, Asaminami, Hiroshima 731-3195, Japan
5. Centre for Marine & Coastal Studies, Universiti Sains Malaysia 11800 USM, Penang, Malaysia
6. Department of Zoology, Graduate School of Science, Kyoto University, Kyoto 606-8502, Japan