1. Department of Biological Sciences, University of Tokyo, Bunkyo-ku, Tokyo 113-0033, Japan
2. Sugashima Marine Biological Laboratory, Graduate School of Science, Nagoya University, Sugashima, Toba, Mie 517-0004, Japan
3. Coastal Branch of Natural History Museum and Institute, Chiba, Kastsuura, Chiba 299-5242, Japan
4. Shimoda Marine Research Center, University of Tsukuba, 5-10-1, Shimoda, Shizuoka 415-0025, Japan