1. Graduate School of Science, Hokkaido University, Sapporo, Hokkaido 060-0810, Japan
2. College of Arts and Sciences, J. F. Oberlin University, Machida, Tokyo 194-0294, Japan
3. Sugashima Marine Biological Laboratory, Graduate School of Science, Nagoya University, Toba 517-0004, Japan
4. Department of Biological Sciences, Graduate School of Science, the University of Tokyo, 7-3-1 Hongo, Bunkyo, Tokyo 113-0033, Japan
5. Graduate School of Sciences and Technology for Innovation, Yamaguchi University, 1677-1 Yoshida, Yamaguchi, Yamaguchi 753-8512, Japan
6. Faculty of Science, Hokkaido University, Sapporo, Hokkaido 060-0810, Japan