1. Shimoda Marine Research Center, University of Tsukuba, 5-10-1 Shimoda, Shizuoka 415-0025, Japan
2. Sugashima Marine Biological Laboratory, Nagoya University, 429-63 Sugashima, Toba, Mie 517-0004, Japan
3. The University Museum, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033, Japan
4. Faculty of Science, Hokkaido University, Kita-ku N10E8, Sapporo, Hokkaido 060-0810, Japan