1. Faculty of Science, University of the Ryukyus, Nishihara, Okinawa 903-0213, Japan
2. School of Regional Innovation and Social Design Engineering, Kitami Institute of Technology, Koen-cho, Kitami, Hokkaido 090-8507, Japan
3. Graduate School of Bioscience, Tokai University, Orido, Shimizu, Shizuoka 424-8610, Japan
4. Center for Marine Environmental Studies, Ehime University, Matsuyama, Ehime 790-8577, Japan
5. School of Marine Science and Technology, Tokai University, Orido, Shimizu, Shizuoka 424-8610, Japan