1. Department of Life Science, International Christian University, 3-10-2 Osawa, Mitaka, Tokyo 181-8585, Japan
2. Graduate School of Arts and Science, International Christian University,3-10-2 Osawa, Mitaka, Tokyo 181-8585, Japan
3. Field Science Center for Northern Biosphere, Hokkaido University, 20-5, Hakodate Research Center for Fisheries and Ocean, Benten-cho, Hakodate, Hokkaido 040-0051, Japan
4. Faculty of Fisheries Science, Hokkaido University, 3-1-1 Minato-cho,Hakodate, Hokkaido 041-8611, Japan