1. Kyoto University, Department of Bioresource Sciences, Kitashirakawa Oiwake-cho, Sakyo-ku, Kyoto 606-8502, Japan
2. Tsuruoka City Kamo Aquarium, Yamagata 997-1206, Japan
3. Kyoto University, Graduate School of Agriculture, Sakyo-ku, Kyoto 606–8502, Japan