1. Faculty of Science, Hokkaido University, Kita-ku N10W8, Sapporo 060-0810, Japan
2. Umusa 774, Nago 905-0006, Japan
3. Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity, Invalidenstraße 43, D-10115, Berlin, Germany
4. Center for Molecular Biodiversity Research, National Museum of Nature and Science, Tsukuba 305-0005, Japan