1. Department of Biological Sciences, Hokkaido University, N10 W8, Sapporo 060-0810, Japan
2. Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK
3. Maine Coastal Program, Department of Marine Resources, 194 McKown Point, West Boothbay Harbor, Maine 04575, USA
4. Scottish Association for Marine Science, Scottish Marine Institute, Oban, Argyll PA37 1QA, Scotland
5. Marine Biological Association, The Laboratory, Citadel Hill, Plymouth PL1 2PB, UK
6. Graduate School of Environmental Science, Hokkaido University, Aikappu 1, Akkeshi, Hokkaido 088-1113, Japan
7. Department of Invertebrate Zoology, Faculty of Biology, Saint Petersburg State University, Universitetskaja nab. 7/9, Saint Petersburg 199034, Russia