1. Department of Applied Science, Kochi University, Kochi 780-8520, Japan
2. Marine Genomics Unit, Okinawa Institute of Science and Technology Graduate University, Onna, Okinawa 904-0495, Japan
3. Kuroshio Science Program, Graduate School of Integrated Arts and Sciences, Kochi University, Kochi 780-8520, Japan
4. College of Ecology and Environment & State Key Laboratory of Marine Resource Utilization in South China Sea, Hainan University, Haikou 570228, China
5. State Key Laboratory of Marine Resource Utilization in South China Sea, Hainan University, Haikou 570228, China
6. Fisheries Technology Institute, Japan Fisheries Research and Education Agency, Ishigaki, Okinawa 907-0451, Japan