1. Nodai Bioresource Institute, Tokyo University of Agriculture, 196 Yasaka, Abashiri, Hokkaido 099-2493, Japan
2. Gene Conservation Laboratory, Alaska Department of Fish and Game, 333 Raspberry Road, Anchorage, Alaska 99518, USA
3. Mariculture Fisheries Research Institute, Hokkaido Research Organization Fisheries Research Department, 1-156-3 Funami-cho, Muroran, Hokkaido 051-0013, Japan
4. Wakkanai Fisheries Research Institute, Hokkaido Research Organization Fisheries Research Department, 4-5-15 Suehiro, Wakkanai, Hokkaido 097-0001, Japan
5. National Research Institute of Far Seas, Fisheries Research Agency, 2-12-4 Fukuura, Kanazawa-ku, Yokohama, Kanagawa 236-8648, Japan
6. Division of Marine Bioscience, Graduate School of Fisheries Sciences, Hokkaido University, 3-1-1 Minato-cho, Hakodate, Hokkaido 041-8611, Japan