1. Misaki Marine Biological Station, School of Science, The University of Tokyo, Misaki, Miura, Kanagawa 238-0225, Japan
2. Bioscience Group, National Institute of Polar Research, Tachikawa, Tokyo 190-0014, Japan
3. Department of Biology, Keio University, Yokohama, Kanagawa 223-8521, Japan
4. Faculty of Environmental Earth Science, Hokkaido University, Sapporo, Hokkaido 060-0810, Japan
5. Animal Evolution and Biodiversity, Johann-Friedrich-Blumenbach Institute for Zoology and Anthropology, Georg-August-Universität Göttingen, Göttingen, Germany