1. Department of Biology, Faculty of Science, Toho University, 2-2-1 Miyama, Funabashi, Chiba 274-8510, Japan
2. Department of Regulatory Biology, Faculty of Sciences, Saitama University, 255 Shimo-okubo, Sakura-ku, Saitama 338-8570, Japan
3. Department of Biology, Faculty of Education and Integrated Arts and Sciences, Center for Advanced Biomedical Sciences, Waseda University, 2-2 Wakamatsu-cho, Shinjyuku-ku, Tokyo 162-8480, Japan