1. Noto Marine Laboratory, Institute of Nature and Environmental Technology, Kanazawa University, Ogi, Noto-cho, Ishikawa 927-0553, Japan
2. Department of Biological Sciences, Faculty of Science, Kanagawa University, 3-27-1, Rokkakubashi, Kanagawa-ku, Yokohama-city, Kanagawa 221-8686, Japan