1. Department of Biology, Graduate School of Science and Technology, Shinshu University, Matsumoto, Nagano 390-8621, Japan
2. Department of Mountain and Environmental Science, Interdisciplinary Graduate School of Science and Technology, Shinshu University, Asahi 3-1-1, Matsumoto, Nagano 390-8621, Japan
3. Department of Biology, Faculty of Science, Shinshu University, Matsumoto, Nagano 390-8621, Japan