1. Laboratory of Functional Biology of Aquaculture, School of Marine Biosciences, Kitasato University, Sagamihara, Kanagawa 252-0373, Japan
2. Department of Physics, School of Science, Kitasato University, Sagamihara, Kanagawa 252-0373, Japan
3. Kitanihon Fishery Co. Ltd., Sanriku, Iwate 022-0211, Japan