1. Tropical Biosphere Research Center, University of the Ryukyus, Okinawa 903-0213, Japan
2. Faculty of Fisheries and Marine Science, Halu Oleo University, Kendari 93232, Indonesia
3. Faculty of Fisheries and Marine Science, Sam Ratulangi University, Manado 95115, Indonesia
4. Graduate School of Medicine, University of the Ryukyus, Okinawa 903-0125, Japan