1. Department of Biological Science and Technology, National Pingtung University of Science and Technology, No.1, Shuefu Road, Neipu, Pingtung 912, Taiwan
2. Department of Forestry and Natural Resources, National Ilan University, No.1, Sec. 1, Shennong Road, Yilan City, Yilan County 260, Taiwan
3. Institute of Wild Life Conservation, National Pingtung University of Science and Technology, No.1, Shuefu Road, Neipu, Pingtung 912, Taiwan