1. Tokyo Sea Life Park, 6-2-3 Rinkai-cho, Edogawa-ku, Tokyo 134-8587, Japan
2. Japan Agency for Marine-Earth Science and Technology (JAMSTEC), 2-15 Natsushima-cho, Yokosuka City, Kanagawa 237-0061, Japan
3. Misaki Marine Biological Station, The University of Tokyo, 1024 Koajiro, Misaki, Miura City, Kanagawa 238-0225, Japan