1. Department of Bioresource Science, Graduate School of Integrated Sciences for Life, Hiroshima University, 1-4-4 Kagamiyama, Higashihiroshima, Hiroshima 739-8528, Japan
2. Fisheries Technology Institute, Japan Fisheries Research and Education Agency, 234 Yashima-higashi, Takamatsu, Kagawa 761-0111, Japan
3. Department of Aquaculture, Bangladesh Agricultural University, Mymensingh 2202, Bangladesh