1. Meguro Parasitological Museum, 4-1-4 Shimomeguro, Meguro, Tokyo 153-0064, Japan
2. Atmosphere and Ocean Research Institute, The University of Tokyo, 5-1-5 Kashiwanoha, Kashiwa, Chiba 277-8564, Japan
3. Misaki Marine Biological Station, Graduate School of Science, The University of Tokyo, 1024 Misaki, Miura, Kanagawa 238-0225, Japan