1. Graduate School of Environmental ScienceHokkaido UniversityKita‐ku060‐0810SapporoJapan
2. Department of ZoologyOxford UniversityNew Radcliffe House, Radcliffe Observatory Quarter, 6GG, Woodstock RdOX2OxfordUK
3. Centre for Biodiversity and Conservation ScienceUniversity of Queensland4071St. LuciaQLDAustralia
4. Evolutionary Demography LaboratoryMax Plank Institute for Demographic Research18057RostockGermany
5. Department of Animal and Plant SciencesUniversity of SheffieldAlfred Denny Building, Western BankS10 2TNSheffieldUK